Family Platycephalidae

Taxonomy: Nelson (2006) includes some 65 valid species of platycephalids in 18 genera. The taxonomy of the family is confused and in need of revision. We treat 44 Australian species within 15 genera.
Distribution, ecology and habitat: The family is widespread primarily in the Indo Pacific, also occurring westward to the Red Sea and southern Africa. Benthic, commonly occurring on mud or sand bottoms from estuaries to about 300m depth on the upper continental slope. The family contains a number of widespread species, as well as localised endemics, further investigation is required to ascertain the status of species complexes and potentially cryptic taxa.
Characteristics:
Body elongate; head moderately to strongly depressed; bony ridges of head frequently bearing sharp spines and serrations; body mostly bearing ctenoid scales, cycloid scales often present ventrally; pelvic fins behind pectoral base; mouth large; lower jaw extending beyond upper when mouth closed. Ocular and interopercular flaps variably developed; eye partly directed upward. Teeth on jaws, vomer and palatines generally caniniform; some species with stout canines. First dorsal fin with VI-X spines; first spine short, isolated or poorly connected to second spine; second dorsal fin with 10-15 soft rays, lacking spines; anal fin lacking spines.
Size: Maximum size greatly variable, from about 10 cm to 120 cm.
Food and feeding: Feed mainly on small fish and crustaceans. Generally ambush predators, will bury in mud with only eyes and upper parts of head exposed, some species are also active foragers.
Reproduction and early life history: Knowledge of the life history and reproduction of flathead varies greatly, species targeted by commercial and recreational fishers are generally well studied, whereas little is known of many non-target species. A number of commercially important species are known to spawn over the warmer months, with mature animals of some species moving to shallow water prior to spawning.
Fisheries: A number of species are regarded as excellent eating and are targeted by recreational and commercial fishers, with several species being important commercially. Commonly caught by line, gillnet and trawl.
Australian species:
- Ambiserrula jugosa (McCulloch, 1914) Mud Flathead CAAB 37296041
- Cociella hutchinsi Knapp, 1996 Brownmargin Flathead CAAB 37296018
- Cymbacephalus bosschei (Bleeker, 1860) Smalleye Flathead CAAB 37296031
- Cymbacephalus nematophthalmus (Günther, 1860) Fringe-eye Flathead CAAB 37296023
- Cymbacephalus staigeri (Castelnau, 1875) Northern Rock Flathead CAAB 37296042
- Elates ransonnetii (Steindachner, 1877) Dwarf Flathead CAAB 37296013
- Inegocia harrisii (McCulloch, 1914) Harris' Flathead CAAB 37296010
- Inegocia japonica (Tilesius, 1812) Rusty Flathead CAAB 37296029
- Kumococius rodericensis (Cuvier, 1829) Whitefin Flathead CAAB 37296019
- Leviprora inops (Jenyns, 1840) Longhead Flathead CAAB 37296005
- Onigocia bimaculata Knapp, Imamura & Sakashita, 2000 Twospot Flathead CAAB 37296050
- Onigocia macrolepis (Bleeker, 1854) Notched Flathead CAAB 37296025
- Onigocia oligolepis (Regan, 1908) Shortsnout Flathead CAAB 37296027
- Onigocia pedimacula (Regan, 1908) Broadband Flathead CAAB 37296051
- Onigocia spinosa (Temminck & Schlegel, 1843) Midget Flathead CAAB 37296022
- Platycephalus arenarius Ramsay & Ogilby, 1886 Northern Sand Flathead CAAB 37296021
- Platycephalus aurimaculatus Knapp, 1987 Toothy Flathead CAAB 37296035
- Platycephalus bassensis Cuvier, 1829 Southern Sand Flathead CAAB 37296003
- Platycephalus caeruleopunctatus McCulloch, 1922 Bluespotted Flathead CAAB 37296007
- Platycephalus chauliodous Knapp, 1991 Bigtooth Flathead CAAB 37296039
- Platycephalus conatus Waite & McCulloch, 1915 Deepwater Flathead CAAB 37296002
- Platycephalus endrachtensis Quoy & Gaimard, 1825 Yellowtail Flathead CAAB 37296020
- Platycephalus fuscus Cuvier, 1829 Dusky Flathead CAAB 37296004
- Platycephalus indicus (Linnaeus, 1758) Bartail Flathead CAAB 37296033
- Platycephalus laevigatus Cuvier, 1829 Rock Flathead CAAB 37296006
- Platycephalus longispinis Macleay, 1884 Longspine Flathead CAAB 37296036
- Platycephalus marmoratus Stead, 1908 Marbled Flathead CAAB 37296038
- Platycephalus richardsoni Castelnau, 1872 Tiger Flathead CAAB 37296001
- Platycephalus speculator Klunzinger, 1872 Southern Bluespotted Flathead CAAB 37296037
- Ratabulus diversidens (McCulloch, 1914) (McCulloch, 1914) CAAB 37296011
- Rogadius asper (Cuvier, 1829) Olive-tail Flathead CAAB 37296024
- Rogadius patriciae Knapp, 1987 Blackbanded Flathead CAAB 37296008
- Rogadius pristiger (Cuvier, 1829) Thorny Flathead CAAB 37296054
- Rogadius serratus (Cuvier, 1829) Serrate Flathead CAAB 37296040
- Sorsogona tuberculata (Cuvier, 1829) Tuberculate Flathead CAAB 37296030
- Suggrundus macracanthus (Bleeker, 1869) Bigspine Flathead CAAB 37296012
- Sunagocia arenicola (Schultz, 1966) Broadhead Flathead CAAB 37296043
- Sunagocia otaitensis (Cuvier, 1829) Fringelip Flathead CAAB 37296046
- Sunagocia sainsburyi Knapp & Imamura, 2004 Sainsbury's Flathead CAAB 37296055
- Thysanophrys celebica (Bleeker, 1854) Celebes Flathead CAAB 37296044
- Thysanophrys chiltonae Schultz, 1966 Longsnout Flathead CAAB 37296034
- Thysanophrys cirronasa (Richardson, 1848) Tasselsnout Flathead CAAB 37296045
- Thysanophrys papillaris Imamura & Knapp, 1999 Smallknob Flathead CAAB 37296053
- Thysanophrys papillolabium Schultz, 1966 Fringe Flathead CAAB 37296047
References:
Eschmeyer, W.N. (2008) The Catalogue of Fishes on-line. (California Academy of Sciences: San Francisco) Available from: http://www.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (12 June, 2008).
Hoese, D.F., Bray, D.J., Paxton, J.R. & Allen, G.R. (2006) Fishes. In: P.L. Beesley & A. Wells (eds) Zoological Catalogue of Australia Volume 35. Parts 1-3. Australian Biological Resources Study and CSIRO Publishing, 2248 pp.
Kailola, P.J., Williams, M.J., Stewart, P.C., Reichelt, R.E., McNee, A. & Grieve, C. (eds) (1993) Australian fisheries resources. Bureau of Resource Sciences, Australia & Fisheries Research and Development Corporation. 422 p.
Knapp, L.W. (1999) Family Platycephalidae, flatheads, pp. 2385-2421. In: K.E. Carpenter & V.H. Niem (eds). FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Volume 4: Bony Fishes Part 2 (Mugilidae to Carangidae). Food and Agriculture Organization of the United Nations, Rome.
Nelson, J.S. (2006). Fishes of the World. 4th Ed. John Wiley & Sons, New York, N.Y.